https://doi.org/10.1351/goldbook.14590
Deviation within a population from a mean of zero for the difference between a trait measured from the right and left sides of bilaterally symmetrical individuals from that population.
Example: Measurement of the difference in muscle mass of left and right arms of humans would display directional asymmetry because most humans are right-handed and have larger right arms.
See also: antisymmetry, fluctuating asymmetry